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As the police arrive at Dexter's house, he is obviously in shock and admits "it was me", feeling guilty that his relationship with Trinity caused her death and/or answering the officers question "sir, did you say that you called this in?". Quinn is already suspicious about the circumstances surrounding Rita's death, considering it was not Trinity's modus operandi. Astor takes the news particularly hard, and blames Dexter for her death. Unable to reconcile, Astor decides that she wants to live with her grandparents in Orlando, and she and Cody leave Miami.
Miami Metro begin investigating a severed head in a Venezuelan neighborhood, and also find several related cases. The suspect is quickly nicknamed the Santa Muerte Killer. The FBI, unable to find Arthur Mitchell, follows its only other lead, Kyle Butler. Quinn recognizes the similarities between sketches of Kyle Butler and Dexter. Dexter finds a Department of Sanitation worker, Boyd Fowler, who is responsible for the deaths of many women. Dexter hires a nanny named Sonya to care for Harrison. Dexter eventually kills Fowler, but the crime is witnessed by Fowler's next victim whom he had in captivity.
Dexter tries to care for the injured woman named Lumen (Julia Stiles), but she is understandably suspicious, asking Dexter if he is going to "sell her". Quinn approaches Jonah Mitchell, who is now in witness protection. He shows Mitchell a picture of Dexter and asks if it is Kyle Butler, but is interrupted by an undercover FBI agent before Mitchell can identify the picture. Quinn is suspended without pay by LaGuerta for disobeying her orders, and she continues to defend Dexter. Quinn seeks the help of a fellow officer to investigate Dexter. With Dexter's help, Deb closes in on the Santa Muerte killer, but he escapes during a hostage stand-off. Lumen tells Dexter that "there were others" with Fowler who did this to her.
Lumen asks Dexter to help her seek revenge, but he initially refuses. After Lumen continues on her own and targets the wrong suspect, Dexter teaches her the importance of knowing a person is guilty. Lumen falsely leads Dexter to believe she leaves Miami. Dexter soon learns that Lumen has remained in Miami after she shoots one of her attackers. Dexter is forced to help her cover it up, moments before Debra arrives to investigate the crime scene. She later reveals to Dexter that killing one of her attackers brought her a sense of peace, and she tearfully recognizes that it will not last, and she will have to find (and kill) the others to find that peace again. Dexter recognizes this as being her own "Dark Passenger", and decides (in part to forgive himself for failing Rita) to help her.
Also, the subplot throughout the season focuses on the relationship of Maria/Angel, who have now come out into the open about their marriage but since the beginning this marriage has been on the rocks. Also it focuses on Debra's growing feelings to Quinn after a one night stand, though she remains unaware that Quinn suspects her brother is Kyle Butler.
Angel gets involved in a bar fight which could cause him to lose his job. Maria saves him by setting up a sting on Agent Liddy (Peter Weller). Liddy develops a friendship with Quinn, both being angry about being "betrayed" by Maria. Quinn has Liddy spy on Dexter.
Meanwhile, Dexter and Lumen hunt the other people responsible for torturing her, including Jordan Chase (Jonny Lee Miller) and people associated with him. As Quinn's relationship with Deb deepens, he tries to call off Liddy, but by this point Liddy has taken pictures of Dexter and Lumen disposing of body parts on Dexter's boat and captured video of them practicing for a kill. Having lost his job, Liddy attempts to blackmail Quinn into busting Dexter, even capturing Dexter. During a struggle, Dexter kills Liddy, but is forced to leave the body behind as Lumen is captured by Jordan Chase, who figured out that she and Dexter were killing his contacts.
Dexter goes to confront Jordan but is called away by Liddy's crime scene, where Quinn is arrested for suspicion of implication in Liddy's death. Later Dexter tracks down where Jordan is holding Lumen captive, and following a brief struggle, Dexter overpowers Jordan, and then allows Lumen to kill Jordan. Deb discovers them, though they are behind transluscent plastic and she is not aware of their identities. Understanding that Lumen must be an escaped victim, Deb sympathizes, and lets them go, not seeing their faces. After Jordan's death, Lumen no longer feels the "Dark Passenger" and she leaves, leaving Dexter to wonder if there is hope for him, even if he is not enthusiastic about it.
Quinn visits Dexter at Harrison's birthday party, thanking him for getting him out of jail, since Dexter clearly faked a blood test to clear Quinn. Quinn and Deb appear to reconcile, and Astor and Cody ask to move in with Dexter for the summer.
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